Joschua Knüppe
Born in 1992 in Mettingen, Germany Began drawing at age 3 2010, diploma (Fachabitur) dicipline design Studying art since 2010 at the Academy for fine Arts Münster Since 2013 in the class of Shana Moulton Since 2014 master student Exhibitions -2012 "Pyrungata", Kunst in der Region, Kloster Gravenhorst -2013 Förderpreisausstellung, Kunsthalle Münster -2013 "Studentennester", Stadtmuseum Münster -2013 "Ausgrabung eines Eurovenator anglicus westfalia", Museumsdorf Detmold -2013 "All Yesterdays", SkF Osnabrück -2014 Förderpreisausstellung, Kunsthalle Münster -2014 "Silvanus" in F24, Münster -2014 "Seeschlangen, schützenswerte Exoten aus den Reiche der Legende", Geomuseum Münster -2015 “Ein lebender Mythos”, Kunstraum Unten, Bochum Scientific work Sachs et al 2015, Cenomanian–Turonian marine amniote remains from the Saxonian Cretaceous Basin of Germany
Illustrations
57 genera
Aetodactylus
This American ornithocheiroid comes from the start of the Late Cretaceous, a time when not much is known from the rocks of the continent.
Wingspan 3 m
Alanqa
Africa's first real azhdarchid discovery was also a neighbor of a few famous dinosaurs like the fish-eating Spinosaurus and the vicious Carcharodontosaurus.
Wingspan 4 m
Alcione
One of the last pterosaurs, Alcione may have been a plunge diver like kingfisher birds.
Wingspan 2 m
Allkaruen
"Ancient brain" from the Jurassic of Argentina helps fill a gap in the pterosaur family tree.

Arcticodactylus
The tiny Trassic pterosaur from Greenland had complex teeth and a varied diet.
Wingspan 24 cm
Argentinadraco
This Argentine pterosaur may have used its blade-like jaws to find food in beach sands.

Arthurdactylus
Arthurdactylus is named for Sir Arthur Conan Doyle, whose novel “The Lost World” envisioned a land of living dinosaurs and pterosaurs on the same Brazilian plateau where its fossils were discovered.
Wingspan 4.6 m
Aymberedactylus
Aymberedactylus is a "chin-crested" tapejarid from the Early Cretaceous Crato Formation lagerstätte of Ceará, northeastern Brazil.

Batrachognathus
This tiny “frog-mouthed” pterosaur was a master hunter of insects during the Late Jurassic in what is now Kazakhstan.
Wingspan 50 cm
Bellubrunnus
The "beautiful one of Brunn" had wingtips that curved forward gracefully, a rather new and hitherto-unseen trait in a pterosaur.
Wingspan 30 cm
Caelestiventus
Caelestiventus is the earliest known pterosaur from the United States.
Wingspan 1.5 m
Changchengopterus
A Tiaojishan wukongopterid, this animal is known from the most tantalizing of remains: A nearly complete juvenile specimen.
Wingspan 47 cm
Chaoyangopterus
They were not the long-necked monster-stork azhdarchids but were probably powerful predators in their own right. Chaoyangopterus gave its name to a whole new breed of terrestrial stalker pterosaur, the likes of which cropped up again and again during the Early Cretaceous.
Wingspan 1.85 m
Cryodrakon
Paleontologists thought the first bones from this giant Canadian pterosaur were from Quetzalcoatlus.
Wingspan 10 m
Cuspicephalus
Cuspicephalus, the largest of the wukongopterids, hails from the Late Jurassic of England. Quite unusual for members of its family, most of which are from China.
Wingspan 1.2 m
Darwinopterus
The greatest revolutionary discovery of pterosaurs in the past decade, Darwinopterus helped determine the evolution of almost the entire family of flying reptiles.
Wingspan 90 cm
Dendrorhynchoides
Anurognathidae This small pterosaur from the Middle Jurassic of China probably filled a similar ecological role as a living swallow: aerial insect hunter.
Wingspan 40 cm
Dorygnathus
This Early Jurassic pterosaur hunted ancient seas with teeth perfect for grasping slippery prey.
Wingspan 1.5 m
Douzhanopterus
This small pterosaur fills an evolutionary gap between early long-tailed pterosaurs and later short-tailed pterosaurs.
Wingspan 75 cm
Feilongus
This double-crested suspension feeder lived in a temperate forest that became one of the world's most productive fossil sites.
Wingspan 2.4 m
Forfexopterus
Scientists have discovered a new flying reptile from the famous Jehol Biota of northeastern China. This new pterosaur has an extremely long narrow skull with teeth modified for filter feeding.

Gegepterus
This jay-sized filter-feeding pterosaur lived alongside famous feathered dinosaurs in a temperate forest in what is now northeastern China.
Wingspan 50 cm
Gladocephaloideus
A pterosaur from the Yixian Formation, this small ctenochasmatid is known from nearly complete remains.
Wingspan 1.1 m
Gnathosaurus
This filter-feeder was one of many Solnhofen pterosaurs, and like many of them it was a small creature that lived close to the sea. It strained its food from the water with a set of long, thin teeth and a rather spoonbill-shaped jaw.
Wingspan 1.7 m
Hateg Island Azhdarchid
This unnamed pterosaur is one of three azhdarchids that acted as top predators on an island in prehistoric Romania.

Hornby Island Azhdarchid
Meet the cat-sized relative of the giraffe-sized azhdarchids that lived on Canada's west coast at the end of the Mesozoic.

Huaxiapterus
A diminutive tapejarid, Huaxiapterus was one of many prolific Jiufotang pterosaurs.
Wingspan 1.5 m
Ian The Wukongopterid
A new but unidentified wukongopterid from the Jurassic Tiaojishan Formation of China suggests that perhaps the males weren't the only ones with the fancy crests.
Wingspan 90 cm
Iberodactylus
This Spanish pterosaur has family ties in China.
Wingspan 4 m
Jidapterus
Chaoyangopterus' kin clearly dominated many parts of the world. The Early Cretaceous world was their oyster. They spread through China and even Brazil, living like their long-necked azhdarchid kin as terrestrial stalkers.
Wingspan 1.7 m
Keresdrakon
Keresdrakon is the second pterosaur discovered at Brazil’s “pterosaur graveyard.”
Wingspan 3 m
Klobiodon
This pterosaur used its teeth to form a cage to trap fish.
Wingspan 1.7 m
Kunpengopterus
The wukongopterids are quite an interesting family of pterosaurs. Little family member Kunpengopterus' claim to fame might be its nearly complete skeleton with a side of regurgitated fish.
Wingspan 90 m
Lacusovagus
Brazil was a hotbed for pterosaurs in the Late Cretaceous. Lacusovagus stood almost as tall as a man and walked over dry ground, snapping up small prey in its mouth.
Wingspan 4.1 m
Liaodactylus
Tiny Liaodactylus is the earliest known filter-feeding pterosaur.
Wingspan 13 cm
Luchibang
The only known specimen of Luchibang had a wingspan of about 2 meters, but was a juvenile when it died, and may have grown much larger.
Wingspan 2 m
Luopterus
Originally thought to be a species of Dendrorhynchoides, the name of this anurognathid honors the late pterosaurologist Lü Junchang.
Wingspan 40 cm
Mistralazhdarcho
This French pterosaur is one of the most completely known European azhdarchids.
Wingspan 5 m
Moganopterus
Filter-feeding pterosaurs don't get bigger than Moganopterus, or even skinnier. It is one of the slimmest pterosaurs around, with a long neck and a spry physique. This animal used its bristly teeth to sift plankton from cool lakes and rivers.
Wingspan 6 m
Mongol Giant
Three neck bones from Mongolia represent one of the largest pterosaurs ever discovered.

Muzquizopteryx
The only known specimen of the nyctosaurid Muzquizopteryx spent years on display in the office of a mine before being described by paleontologists.
Wingspan 2 m
Mythunga
100 million years ago, Mythunga, a giant fish-eating pterosaur, soared over Australian seas.
Wingspan 4.7 m
Normannognathus
This Jurassic pterosaur had a tall crest on its snout shaped like a breaking wave.

Ordosipterus
The strong jaws and blunt teeth of Ordosipterus helped it break the shells of its food.
Wingspan 3 m
Palaeocursornis
Once thought to be the earliest ratite bird, Palaeocursornis was probably an early azhdarchoid pterosaur instead.
Wingspan 75 cm
Pangupterus
The newest entry into the ranks of the flying reptiles, Pangupterus liui also had the oddest set of teeth of any pterosaur.
Wingspan 70 cm
Parapsicephalus
This early pterosaur is named for its discoverer, Reverend D. W. Purdon.
Wingspan 1.8 m
Phosphatodraco
This azhdarchid hunted in arid northern Africa in the final one million years of the reign of the pterosaurs.
Wingspan 5 m
Quetzalcoatlus
The tallest flying animal ever, and often seen sharing the title of "Largest Flying Vertebrate" with Hatzegopteryx, this was also one of the last of its kind. Quetzalcoatlus soared over North America, meeting the famous likes of T. rex and Triceratops before vanishing like the rest of its neighbors 66 million years ago.
Wingspan 10 m
Sericipterus
This Jurassic pterosaur is one of the largest known long-tailed forms.
Wingspan 1.75 m
Shenzhoupterus
Another Jiufotang pterosaur, Shenzhoupterus is named after an old word for China, "Shenzhou," meaning "divine land."
Wingspan 1.4 m
Tethydraco
Tethydraco fished in the Atlantic Ocean less than a million years before an asteroid caused the K-Pg Extinction.
Wingspan 5 m
Uktenadactylus
Found in the vicinity of Fort Worth, Texas, USA, Uktenadactylus was a large fishing pterosaur.
Wingspan 4 m
Vesperopterylus
This small insectivore may have been able to perch on tree branches.

Wukongopterus
The type genus of the Wukongopteridae, this was the pterosaur that gave its name to the entire family of oddball flyers.
Wingspan 73 cm
Xericeps
Known from just a jaw fragment, Xericeps offers rare insight into continental pterosaurs from Africa.

Yixianopterus
Even though the only known specimen is incomplete and partially forged, Yixianopterus is one of the best-known lonchodectids.
Wingspan 1.8 m
